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About | Classical Genetics | Timelines | What's New | What's Hot

About | Classical Genetics | Timelines | What's New | What's Hot


The Electronic Scholarly Publishing Project: Providing world-wide, free access to classic scientific papers and other scholarly materials, since 1993.


ESP Links

Other Useful Sites

The Electronic Scholarly Publishing Project offers links to the following to other sites that may be of interest to the ESP community. Every effort is made to ensure the relevance of the material and the validity of the links, but the internet is dynamic and things may have changed since these materials were last updated.

To report broken links, inappropriate content, or to make suggestions for new links to be added, CLICK HERE.

MendelWeb is an educational resource for teachers and students interested in the origins of classical genetics, introductory data analysis, elementary plant science, and the history and literature of science. Constructed around Gregor Mendel's 1865 paper "Versuche über Pflanzen-Hybriden" and a revised version of the English translation by C. T. Druery and William Bateson, "Experiments in Plant Hybridization", MendelWeb is offered as a public sourcebook and collaborative environment compatible with a variety of guided and independent studies.
The Mendel Newsletter is an annual publication of the Library of the American Philosophical Society promoting awareness of archival resources for the history of genetics and allied sciences.
The Biology Project is an interactive online resource for learning biology developed at The University of Arizona. The Biology Project is fun, richly illustrated, and tested on 1000s of students. It has been designed for biology students at the college and high school level, but is useful for medical students, physicians, science writers, and all types of interested people. It includes a page dedicated to Mendelian Genetics.
The Dolan DNA Learning Center (DNALC) is the world's first science center devoted entirely to genetics education and is an operating unit of Cold Spring Harbor Laboratory, an important center for molecular genetics research. The DNALC, DNALC West, and Harlem DNA Lab extend the Laboratory's traditional research and postgraduate education mission to the college, precollege, and public levels.
Flybase offers a Database of Drosophila Genes & Genomes. Beginning with the work of Morgan's group at Columbia in the early 20th century, Drosophila has been a workhorse for genetics research. Drosophila The FlyBase project is carried out by a consortium of Drosophila researchers and computer scientists at: Harvard University, University of Cambridge (UK), Indiana University and the University of New Mexico.
Project Gutenberg is the first and largest single collection of free electronic books, or eBooks. Michael Hart, founder of Project Gutenberg, invented eBooks in 1971 and continues to inspire the creation of eBooks and related technologies today.
The Journal of Electronic Publishing (JEP) is a forum for research and discussion about contemporary publishing practices, and the impact of those practices upon users. Our contributors and readers are publishers, scholars, librarians, journalists, students, technologists, attorneys, retailers, and others with an interest in the methods and means of contemporary publishing. At its inception in January 1995, JEP carved out an important niche by recognizing that print communication was in the throes of significant change, and that digital communication would become an important — and in some cases predominant — means for transmitting published information.
As the leading ePublishing platform, HighWire Press partners with independent scholarly publishers, societies, associations, and university presses to facilitate the digital dissemination of 1334 journals, reference works, books, and proceedings. HighWire also offers a complete manuscript submission, tracking, peer review, and publishing system for journal editors, Bench>Press. HighWire provides outstanding technology and support services, and fosters a dynamic and innovative community, enhancing the strengths of each of its members.

HighWire plays a unique role in the online publishing business, as part of an academic library, a first-class research institution, and an extension of the publishing world. HighWire is a partner and colleague to scholarly societies who value the quality and content of their journals and other materials. By working with HighWire, publishers gain the economies of scale and networking opportunities of working collaboratively in their business technology.
The National Science Digital Library is a national network dedicated to advancing STEM (Science, Technology, Engineering and Mathematics) teaching and learning for all learners and both formal and informal settings, and the locus of activity for the National Science Foundation's National STEM Distributed Learning program. NSDL receives the majority of its funding through the generous support of NSF's Division of Undergraduate Education (DUE), Directorate for Education and Human Resources (EHR).
Google Scholar provides a simple way to broadly search for scholarly literature. From one place, you can search across many disciplines and sources: articles, theses, books, abstracts and court opinions, from academic publishers, professional societies, online repositories, universities and other web sites. Google Scholar helps you find relevant work across the world of scholarly research.
Evolution was developed by Donald Forsdyke (of Queen's University, Kingston, Ontario, Canada) to provide original works and contextual commentary on matters relating to evolutionary thinking.
ECHO (Exploring and Collecting History Online) is a directory to 5,000+ websites concerning the history of science, technology, and industry. You can search it, browse it according to category, or even look at the tag cloud we've generated. Every website contains a brief description (some examples are here), and occasionally, a review (like these).
Online since 2002, The Complete Work of Charles Darwin Online (or Darwin Online) is the largest and most widely consulted edition of the writings of Darwin ever published. More copies of Darwin's works have been downloaded from Darwin Online than have been printed by all publishers of the past 180 years combined.

This website contains over 91,000 pages of searchable text and 209,000 electronic images, at least one exemplar of all known Darwin publications, reproduced to the highest scholarly standards, both as searchable text and electronic images of the originals. The majority of these have been edited and annotated for the first time with thousands of original editorial notes.

This website also provides the largest collection of Darwin's private papers and manuscripts ever published: c. 20,000 items in c. 100,000 images, thanks primarily to the kind permission of Cambridge University Library. Thus Darwin Online makes available not only Darwin's published science, but the notes and data collected to create it.
GALTON.ORG is edited and maintained by Gavan Tredoux on a voluntary basis. All material provided on this site is made available in good faith for educational and scholarly purposes only. The intention of the editor is to make everything that Galton wrote available here, usually in facsimile format. The site now runs to some 2 gigabytes of digitized material.
The American Museum of Natural History is one of the world's preeminent scientific and cultural institutions. Since its founding in 1869, the Museum has advanced its global mission to discover, interpret and disseminate information about human cultures, the natural world and the universe through a wide-ranging program of scientific research, education and exhibition. The Museum is renowned for its exhibitions and scientific collections, which serve as a field guide to the entire planet and present a panorama of the world's cultures.
The Smithsonian Institution in Washington DC is the world's largest museum complex and research organization. It is composed of 19 museums, 9 research centers, and the National Zoo.

ESP Quick Facts

ESP Origins

In the early 1990's, Robert Robbins was a faculty member at Johns Hopkins, where he directed the informatics core of GDB — the human gene-mapping database of the international human genome project. To share papers with colleagues around the world, he set up a small paper-sharing section on his personal web page. This small project evolved into The Electronic Scholarly Publishing Project.

ESP Support

In 1995, Robbins became the VP/IT of the Fred Hutchinson Cancer Research Center in Seattle, WA. Soon after arriving in Seattle, Robbins secured funding, through the ELSI component of the US Human Genome Project, to create the original ESP.ORG web site, with the formal goal of providing free, world-wide access to the literature of classical genetics.

ESP Rationale

Although the methods of molecular biology can seem almost magical to the uninitiated, the original techniques of classical genetics are readily appreciated by one and all: cross individuals that differ in some inherited trait, collect all of the progeny, score their attributes, and propose mechanisms to explain the patterns of inheritance observed.

ESP Goal

In reading the early works of classical genetics, one is drawn, almost inexorably, into ever more complex models, until molecular explanations begin to seem both necessary and natural. At that point, the tools for understanding genome research are at hand. Assisting readers reach this point was the original goal of The Electronic Scholarly Publishing Project.

ESP Usage

Usage of the site grew rapidly and has remained high. Faculty began to use the site for their assigned readings. Other on-line publishers, ranging from The New York Times to Nature referenced ESP materials in their own publications. Nobel laureates (e.g., Joshua Lederberg) regularly used the site and even wrote to suggest changes and improvements.

ESP Content

When the site began, no journals were making their early content available in digital format. As a result, ESP was obliged to digitize classic literature before it could be made available. For many important papers — such as Mendel's original paper or the first genetic map — ESP had to produce entirely new typeset versions of the works, if they were to be available in a high-quality format.

ESP Help

Early support from the DOE component of the Human Genome Project was critically important for getting the ESP project on a firm foundation. Since that funding ended (nearly 20 years ago), the project has been operated as a purely volunteer effort. Anyone wishing to assist in these efforts should send an email to Robbins.

ESP Plans

With the development of methods for adding typeset side notes to PDF files, the ESP project now plans to add annotated versions of some classical papers to its holdings. We also plan to add new reference and pedagogical material. We have already started providing regularly updated, comprehensive bibliographies to the ESP.ORG site.

Electronic Scholarly Publishing
21454 NE 143rd Street
Woodinville, WA 98077

E-mail: RJR8222 @

Papers in Classical Genetics

The ESP began as an effort to share a handful of key papers from the early days of classical genetics. Now the collection has grown to include hundreds of papers, in full-text format.

Digital Books

Along with papers on classical genetics, ESP offers a collection of full-text digital books, including many works by Darwin (and even a collection of poetry — Chicago Poems by Carl Sandburg).


ESP now offers a much improved and expanded collection of timelines, designed to give the user choice over subject matter and dates.


Biographical information about many key scientists.

Selected Bibliographies

Bibliographies on several topics of potential interest to the ESP community are now being automatically maintained and generated on the ESP site.

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ESP Picks from Around the Web (updated 11 MAY 2018 )